I found the following from the Nikon support website, hope it helps.
Use one or multiple extension rings for further creative capabilities. An AI NIKKOR will retain automatic diaphragm and meter coupling functions, letting you compose, focus and meter at a wide-open apertures. The rings can also be used for manual exposure control (Exposure meter does not work with D90, D5100, D5000, D3100 and D3000).
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