VR is Nikons term for vibration reduction, or image stabilisation. Lenses that have 'VR' can be used at a slower shutter speed than non VR lenses, but only when shooting stationary subjects.
When shooting with a long focal length lens, say 300mm, when you look through your cameras viewfinder the image you see will be jumping about and vibrating due to your inability to hold the camera perfectly still. With a VR lens, when you press the shutter button to focus the image will steady up as the VR system moves a lens element that counteracts your own movements, this results in a pretty steady image and will allow you to use a slower shutter speed than usual